In 2268, Commodore Wesley commanded the USS Lexington in the disastrous M-5 war games. During that tactical exercise, the M-5 computer malfunctioned and the USS Enterprise was deemed out-of-control, following the destruction of the USS Excalibur . Wesley requested permission to destroy the Enterprise with his remaining battle force, to end the threat that the rogue ship posed. His request was approved. However, fortunately, the Enterprise was able to deactivate the M-5 computer and drop its shields before Wesley's battle force could attack. For his part, although communication with the drifting ship was impossible at that point, Wesley successfully gambled that Kirk had gotten the situation under control and held his fire to confirm that conjecture. (TOS: "The Ultimate Computer")
Wesley retired from Starfleet to become governor of Mantilles, the most remote inhabited planet in the Federation. The planet was threatened by a cosmic cloud in 2269, but Governor Wesley remained calm and prevented panic from spreading on Mantilles. He evacuated 5,000 children from Mantilles, in an attempt to spare their lives. Their ships were later recalled, following the diversion of the cosmic cloud. (TAS: "One of Our Planets Is Missing")
Wesley, who was born in Limerick, Ireland in 2216. He had a wife, Fiona, and three sons, Robert Jr., Bruce and Roger, all of whom lived in Pittsburgh, Pennsylvania. His interests included team sports, waterskiing, xenozoology, jazz piano.
Wesley graduated from the Starfleet Academy class of 2238, and had since earned fourteen citations for bravery, 2238-50; the Grankite Order of Tactics (Class of Excellence), 2242; Prentares Ribbon of Commendation, 2243; Karagite Order of Heroism, 2247; Palm Leaf of Lyceus, 2251; Silver Palm With Cluster, 2254; Medal of Valor, 2258.
Wesley's previous assignments included:
- 2238-41: USS Glasgow, under the command of Captain Hart, where he served as junior tactical officer.
- 2241-45: USS Sulaco, under the command of Captain Hicks, where he served as chief tactical officer. Participated in the Battle of Donatu V in 2242, and later became promoted to lieutenant commander, and second officer, following the loss of several crewmembers during the battle.
- 2245-50: USS Enterprise, under the command of Captain Robert April, where he served as first officer.
- 2251-62: Promoted to captain of the USS Beowulf.
- 2263-68: Promoted to commodore and given the command of the USS Constitution.
- 2268: Transferred to command of the USS Lexington for war-games exercise.