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The memory loss fail-safe system was a defense built into the programming of Lieutenant Commander Data. In the event of a memory loss, it was intended to protect the android from anyone who might try to take advantage of him.
The system was activated in 2375 when some of Data's memory engrams, located within his positronic matrix, were damaged by a Son'a weapon on the Ba'ku planet. (Star Trek Nemesis; Star Trek: Insurrection)