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Memory Alpha

The Leyron were a species native to the planet Malkus IX.

They were considered unique as they were one of few known species to have developed a written language significantly before developing a spoken or sign language. (TNG: "Loud As A Whisper")

Kathryn Janeway once studied the gestural idioms of the Leyron, which she found more comprehensible than the non-verbal language of the Tak Tak. (VOY: "Macrocosm")

This species was only mentioned in dialogue.
According to the Star Trek Encyclopedia (4th ed., vol. 2, p. 7), the sign language M-9 was developed by the Leyrons.
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