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[[File:Klingon augment virus Levodian flu.jpg|thumb|The Klingon augment virus]]
The '''Klingon augment [[virus]]''' was a modified form of [[Levodian flu]] that threatened to wipe out the [[Klingon]] race in the mid-[[22nd century]]. It was inadvertently created by Klingon researchers who were attempting to bio-engineer enhanced warriors using [[DNA]] from genetically-modified human [[embryo]]s left over from [[Earth]]'s [[Eugenics Wars]].
 
  +
{{aquote|We do not discuss it with outsiders.|Worf|2268|Trials and Tribble-ations}}
 
The '''Klingon augment [[virus]]''' was a modified form of [[Levodian flu]] that threatened to wipe out the [[Klingon]] race in the mid-[[22nd century]]. It was inadvertently created by Klingon [[researcher]]s who were attempting to bio-engineer enhanced warriors using [[DNA]] from [[genetic engineering|genetically-modified]] [[Human]] [[embryo]]s left over from [[Earth]]'s [[Eugenics Wars]].
   
 
== History ==
 
== History ==
 
[[File:Klingon cranial ridges dissolve.jpg|thumb|left|Dissolving cranial ridges]]
 
[[File:Klingon cranial ridges dissolve.jpg|thumb|left|Dissolving cranial ridges]]
In [[2154]], Klingon [[scientist]]s working under the direction of [[Antaak]] recovered some [[augment]] DNA from a destroyed [[Klingon Bird-of-Prey (22nd century)|Bird of Prey]] that had earlier been hijacked by [[Human]] augments created by [[Arik Soong]]. Fearful that Humans would create augments and overwhelm the [[Klingon Empire|Empire]], the scientists attempted to create their own augments to counter the perceived threat.
+
In [[2154]], Klingon [[scientist]]s working under the direction of [[Antaak]] recovered several [[Augment]] embryos from a hijacked [[Klingon Bird-of-Prey (22nd century)|Bird-of-Prey]] destroyed in orbit of [[Qu'Vat colony]] during the [[Augment Crisis]]. Fearing that Humans could overwhelm the [[Klingon Empire|Empire]] if they moved to staffing their ships with Augments, a project was set up to attempt to create their own Augments to counter the perceived threat.
   
The effort failed dreadfully. The augments created did develop enhanced strength and intelligence, but they also began to show Human characteristics, from the blatant to the subtle. These included personality alterations and loss of the characteristic forehead ridges. Ultimately, the subjects died agonizing deaths when the incompatible DNA resulted in neural system breakdown. While working to stabilize the process of augmentation, the scientific team augmented a test subject who was, unknown to the team, also infected with Levodian flu. The flu virus in some manner incorporated the augment DNA, turning a carefully controlled experiment into an epidemic. In as much as the unstable augmentation process always killed the subject, this epidemic threatened the Klingon race with extinction.
+
The effort failed dreadfully. The Augments created did develop enhanced strength and intelligence, but they also began to show Human characteristics, from the blatant to the subtle. These included personality alterations and loss of the characteristic forehead ridges. Ultimately, the subjects died agonizing deaths when the incompatible DNA resulted in neural system breakdown. While working to stabilize the augmentation process, the scientific team enhanced a test subject who was, unknown to the team, also infected with Levodian flu. The flu virus in some manner incorporated the Augment DNA, turning a carefully controlled experiment into an epidemic. As the unstable augmentation process always killed the subject, this epidemic threatened the Klingon race with [[extinction]].
   
  +
The virus spread very rapidly, infecting millions within months. One of the first planets struck was [[N'Vak colony]]. The Klingons attempted to capture [[Doctor]] [[Arik Soong]] to help develop a cure, but their efforts were thwarted by Soong's incarceration in a high security detention facility. They then moved to [[Kidnapping|kidnap]] Doctor [[Phlox]], using [[Rigelian]] intermediaries and the assistance of [[Section 31]] agents. Phlox was forced to work with Antaak to find a cure. However, the [[Klingon High Council]] was unwilling to wait, and dispatched a fleet to annihilate the infected planets, starting with N'Vak colony. Afterward it set course to Qu'Vat colony. ({{ENT|Affliction}})
 
[[File:Antaak.jpg|thumb|Antaak before]]
 
[[File:Antaak.jpg|thumb|Antaak before]]
 
[[File:Antaak-ridgeless.jpg|thumb|Antaak after]]
 
[[File:Antaak-ridgeless.jpg|thumb|Antaak after]]
The Klingons attempted to capture Arik Soong to help them develop a cure, but their efforts were thwarted by Soong's high security detention. The Klingons next arranged to kidnap [[Doctor]] [[Phlox]] using [[Rigelian]] intermediaries and with the assistance of [[Section 31]] agents. Phlox was forced to work with Antaak to stabilize the augment virus. Antaak's goal was to create a stable method of creating Klingon augments, while Phlox's goal was to cure the plague before the Klingons died as a species. Ultimately, the goal achieved lay somewhere in the middle: a method was created that stabilized the process in the early stages – after the cranial ridges dissolved and some minor neural re-ordering had occurred (which caused personality changes), but before the fatal [[neural pathway]] breakdown. Because afflicted Klingons' DNA had been altered by the virus, these traits were passed onto their descendants. ({{ENT|Affliction|Divergence}})
 
   
  +
When it became clear to the research team on Qu'vat colony that it would take weeks to develop a cure, stabilizing the augment virus became the researcher's primary concern. Their efforts proved successful; a method was developed that stabilized the process in the early stages, after the cranial ridges had dissolved and some minor neural re-ordering had occurred (which caused personality changes), but before the virus became contagious, enhanced augment abilities, and caused a fatal [[neural pathway]] breakdown.
Evidently, the [[Excalbian]]s only knew of affected Klingons, as they depicted [[Kahless]] himself as affected. This may, however, have been attributed to reading the minds of humans who, at that point in history, only had dealings with Klingon augments. ({{TOS|The Savage Curtain}})
 
   
  +
Eventually, [[Admiral]] {{dis|Krell|22nd century}} convinced the High Council to call off their sterilization program of destroying the infected planets. The cure was distributed throughout the Empire. Because afflicted Klingons' DNA had been altered by the virus, the Human appearance and genetic traits were passed onto the descendants of the infected. Initially there were millions of Klingons who had to live with the disfigurement. ({{ENT|Divergence}})
It would be over a century before they discovered a method of restoring all Klingons to their proper appearance. It is unknown if all the damage done has been reversed, evidenced by the change in color to Klingon [[blood]]. ({{film|1}}, {{film|6}}, ''[[Star Trek: The Next Generation]]'')
 
   
  +
[[File:Klingon invasion of Organia.jpg|thumb|A group of Klingons descended from those infected with the augment virus in [[2267]]]]
The Klingons were apparently so embarrassed by the fallout from their failed attempt at genetic enhancement that they refused to discuss the incident with outsiders. Due to the secrecy of the Klingon Empire, knowledge of the change became lost over time to the general population of the Federation. By the [[24th century]], the reason for Klingons having smooth foreheads was not widely known outside the Empire, and questions were generally met with a brusque answer along the lines of "we don't discuss it with outsiders". ({{DS9|Trials and Tribble-ations}})
 
  +
In 2154, Antaak and [[Laneth]] feared that Klingons without ridges would become outcasts, lose their positions and would no longer have a place in the Empire. ({{ENT|Divergence}}) However, by the [[2260s]], affected Klingons served as the [[captain]]s and crews of front-line starships in the [[Klingon Defense Force]]. (''[[Star Trek: The Original Series]]'') Later on, some of them even achieved the title of [[Dahar Master]] and earned their statues among the [[Hall of Heroes]] on [[Qo'noS]]. ({{DS9|Blood Oath|The Sword of Kahless}})
   
  +
By the [[24th century]], knowledge of Klingons with smooth foreheads had been generally forgotten by the population of the {{Federation}}. Also, the reason for Klingons having smooth foreheads a hundred years earlier was not widely known outside the Empire. When asked by [[Miles O'Brien]] and [[Julian Bashir]], [[Worf]] simply gave the answer "we don't discuss it with outsiders", though Bashir immediately suspected viral mutation to be the cause. ({{DS9|Trials and Tribble-ations}})
== Background Information ==
 
This storyline, seen in the ''Enterprise'' episodes {{e|Affliction}} and {{e|Divergence}}, was intended in part to explain why [[Star Trek: The Original Series|''Original Series'']] Klingons had a more human-like appearance, and later Klingons did not. The writers also hoped the two-parter would shed some light on the change in the Klingons' temperament and disposition between the ''TOS'' and ''TNG'' eras.
 
   
  +
{{bginfo|In {{TOS|The Savage Curtain}}, the [[Excalbian]]s created [[Kahless]] as an affected Klingon. This may however be attributed to the fact that they created him from the minds of Humans who at that point mostly had dealings with smooth forehead Klingons.|A cure to restore Klingons to their proper appearance was presumably found some time between [[2268]] and [[2293]], as [[Kang]] was seen as a smooth-headed Klingon in {{TOS|Day of the Dove}} and as a normal Klingon in {{VOY|Flashback}}. The first time normal Klingons were seen on screen was in {{film|1}} set in the early-[[2270s]].}}
For years, unofficial explanations had appeared in ''Star Trek'' literature to explain the difference between the Klingons from the 1960s series and those in later productions. The idea of genetic engineering was explored heavily in several publications licensed by Paramount Pictures, among them the [[Star_Trek_Roleplaying_Game | Roleplaying Game]] published by [[FASA]], several [[Pocket Books]] novels (such as "[[Rules of Engagement]]"), and the reference book ''"[[The Worlds of the Federation]]"'' by author [[Shane Johnson]]. These works indicated that the "Klingons" encountered during the Original Series era were "Klingon-Human fusions" intentionally created to make infilitration into Federation space easier. The true nature of Klingons was revealed during the emergency transmission of the {{IKS|Amar}} as depicted in {{film|1}}.
 
   
 
== Background information ==
In addition to the "fusion" explanation, the producers of ''Star Trek: Enterprise'' had heard many pitches over the years for stories aimed at explaining the change in the Klingon appearance. Additional theories included that the Klingons seen during the Original Series were from another race than those of later eras and also that some sort of disease was to blame. The ''Enterprise'' writing staff mirrored the fan community in that some felt it might be fun to account for the change, while others thought it was probably best to ignore the entire situation. The topic came up again when writer [[Manny Coto]] was named showrunner during the series' [[ENT Season 4|fourth season]].
 
 
This storyline, seen in the ''Enterprise'' episodes {{e|Affliction}} and {{e|Divergence}}, was intended in part to explain why ''[[Star Trek: The Original Series|Original Series]]'' Klingons had a more Human appearance, and later Klingons did not. The writers also hoped the two-parter would shed some light on the change in the Klingons' temperament and disposition between the TOS and TNG eras.
   
 
For years, unofficial explanations had appeared in ''Star Trek'' literature to explain the difference between the Klingons from the 1960s series and those in later productions. The idea of genetic engineering was explored heavily in several publications licensed by Paramount Pictures, among them [[FASA]]'s ''[[Star Trek: The Role Playing Game (FASA)|Star Trek: The Roleplaying Game]]'', several [[Pocket Books]] novels (such as ''[[Rules of Engagement]]''), and the reference book ''[[The Worlds of the Federation]]'' by author [[Shane Johnson]]. These works indicated that the "Klingons" encountered during the ''Original Series'' era were "Klingon-Human fusions" intentionally created to make infiltration into Federation space easier. The true nature of Klingons was revealed during the emergency transmission of the {{IKS|Amar}} as depicted in {{film|1}}.
At first, the producers were only interested in a story with one or more "ridgeless" Klingons who had infiltrated Starfleet for the purpose of intelligence gathering. It was thought the story might involve a surgically-altered Klingon operative aboard the ''Enterprise,'' someone like [[Arne Darvin]] a century later. Around this time, the writing staff had recently concluded the three-part [[Augment Crisis]] arc, and it occurred to them that some of the genetically-engineered embryos might have survived the destruction of the ''Bird of Prey'', and that the Klingons might use these embryos to bio-engineer their own version of "Klingon supermen". This seemed to be a way into a story dealing with the origin of human-like Klingons. More to the point, the ''Enterprise'' producers thought it was simply "too cool" an idea to reveal that [[Kor]], [[Kang]] and other ''Original Series'' Klingons may have had the DNA of [[Khan Noonien Singh|Khan]] inside them.
 
   
 
In addition to the "fusion" explanation, the producers of ''Star Trek: Enterprise'' had heard many pitches over the years for stories aimed at explaining the change in the Klingon appearance. Additional theories included that the Klingons seen during the ''Original Series'' were from another race than those of later eras and also that some sort of disease was to blame. The ''Enterprise'' writing staff mirrored the fan community in that some felt it might be fun to account for the change, while others thought it was probably best to ignore the entire situation. The topic came up again when writer [[Manny Coto]] was named showrunner during the series' [[ENT Season 4|fourth season]].
While some fans clamored for an explanation for the "smooth forehead" Klingons, "Affliction" teleplay writer [[Mike Sussman]] hoped to do more account for the apparent change in Klingon culture between the 23rd and 24th centuries. As depicted in the first ''Star Trek'' series, Klingons were notoriously savage, crafty and at times even cowardly, putting this characterization at odds with the more noble and honorable Klingons seen in ''The Next Generation''. The "minor neural re-ordering" noted by Phlox was intended to suggest that the augmented Klingons were not only "cursed" with a more human appearance, but that they also inherited many human weaknesses as well. In "Divergence", the female [[Laneth]], newly infected by the Augment virus, claimed that she "felt fear for the first time since I was a child." Her fellow augmented warriors had "become like (humans)... weak, cowardly."
 
   
 
At first, the producers were only interested in a story with one or more "ridgeless" Klingons who had infiltrated Starfleet for the purpose of intelligence gathering. It was thought the story might involve a surgically-altered Klingon operative aboard the ''Enterprise,'' someone like [[Arne Darvin]] a century later. Around this time, the writing staff had recently concluded the three-part [[Augment Crisis]] arc, and it occurred to them that some of the genetically-engineered embryos might have survived the destruction of the Bird-of-Prey, and that the Klingons might use these embryos to bio-engineer their own version of "Klingon supermen". This seemed to be a way into a story dealing with the origin of Human-like Klingons. More to the point, the ''Enterprise'' producers thought it was simply "too cool" an idea to reveal that [[Kor]], [[Kang]], and other ''Original Series'' Klingons may have had the DNA of [[Khan Noonien Singh|Khan]] inside them.
The ''Enterprise'' writers' explanation for the change in the Klingons did not and could not satisfy every ''Star Trek'' fan. [[Gene Roddenberry]] himself reportedly believed any "explanation" was unnecessary; the makeup seen in the films and the later series would have been too expensive during the 1960s. Roddenberry felt it was best to simply imagine that Klingons always had ridges (although this preference was perhaps tossed out the window when the change was noted by the DS9 crew in {{e|Trials and Tribble-ations}}). Still, the ''Enterprise'' writers felt that the "dramatic convention" explanation '''was and is still valid''', if a viewer prefers it. If one accepts Roddenberry's suggestion that TOS-era Klingons always had ridges – and that the DS9 reference was merely a joke by the episode writers – then perhaps the Augment virus had no lasting effect on the Empire: the disease may have been cured relatively quickly following the events seen in the ''Enterprise'' two-parter. After all, there was no reference to it in the "later" series (which of course were produced years ''before'' ''[[Star Trek: Enterprise]]'').
 
  +
{{bginfo|This background information was contributed in part by former ''Enterprise'' writer/producer [[User:Mdsussman|Mike Sussman]], a member of Memory Alpha.}}
 
 
While some fans clamored for an explanation for the "smooth forehead" Klingons, "Affliction" teleplay writer [[Mike Sussman]] hoped to do more – account for the apparent change in Klingon culture between the 23rd and 24th centuries. As depicted in the first ''Star Trek'' series, Klingons were notoriously savage, crafty, and at times even cowardly, putting this characterization at odds with the more noble and honorable Klingons seen in ''The Next Generation''. The "minor neural re-ordering" noted by Phlox was intended to suggest that the augmented Klingons were not only "cursed" with a more Human appearance, but that they also inherited many Human weaknesses as well. In "Divergence", the female [[Laneth]], newly infected by the augment virus, claimed that she "felt fear for the first time since I was a child." Her fellow augmented warriors had "become like (Humans)... weak, cowardly."
  +
 
The ''Enterprise'' writers' explanation for the change in the Klingons did not – and could not – satisfy every ''Star Trek'' fan. [[Gene Roddenberry]] himself reportedly believed any "explanation" was unnecessary; the makeup seen in the films and the later series would have been too expensive during the 1960s. Roddenberry felt it was best to simply imagine that Klingons always had ridges (although this preference had already been contradicted by canon when the change was noted by the DS9 crew in {{e|Trials and Tribble-ations}}).
 
{{bginfo|This background information was contributed in part by former ''Enterprise'' writer/producer [[User:Mdsussman|Mike Sussman]], a member of the Memory Alpha community.}}
   
 
== Apocrypha ==
 
== Apocrypha ==
  +
In ''[[The IDIC Epidemic]]'', an original series novel written during the airing of ''The Next Generation'', a Klingon scientist, Korsal, refers to the "Imperial Race," a reclusive breed of Klingons who retained their brow ridges. The Imperial Race has never been seen outside the Empire and its existence is largely a mystery to the Federation. The plague that is the subject of the novel is proven to be a mutated form of "The Imperial Plague," to which the Imperial race is very vulnerable, while smooth-headed Klingons are immune. This novel contained its own partial explanation for the difference in appearance between Klingons from the ''Original Series'' and ''The Next Generation - ''to wit, that "smooth-headed" Klingons were an offshoot of the Klingon race specifically engineered or selected for contact with outside forces.
In "[[Against Their Nature]]", the first installment of the [[IDW Comics]] "[[Klingons: Blood Will Tell]]" series, it is suggested that, while Phlox's cure removed Augment strength and Augment intelligence, those affected did retain Augment ambition, and as such they were largely responsible for the expansion of the Klingon Empire between ''[[Star Trek: Enterprise|Enterprise]]'' and [[TOS]], and were able to take control of the High Council.
 
  +
 
In "[[Against Their Nature]]", the first installment of the [[IDW Comics]] ''[[Star Trek: Klingons - Blood Will Tell]]'' series, it is suggested that, while Phlox's cure removed Augment strength and Augment intelligence, those affected did retain Augment ambition, and as such they were largely responsible for the expansion of the Klingon Empire between ''[[Star Trek: Enterprise|Enterprise]]'' and [[TOS]], and were able to take control of the High Council.
   
In the novel ''[[Forged in Fire]]'', it is revealed that the virus did not solely effect Klingons; the ridged [[Trill]] witnessed in "[[The Host (episode)|The Host]]"- as opposed to the spotted Trill seen in later episodes- are the result of a strain of the Augment virus that managed to infect a Trill colony through visiting Klingon traders. However, the Trill seemed not to be interested in working on methods of restoring the original Trill look, with this 'sub-group' having recently (at the time the novel was set) being re-accepted back into Trill society. The novel also reveals that the appearance of [[The Albino]] - who is not only Klingon but is in fact Kor's cousin - was an unintended side effect of pre-natal genetic engineering intended to cure his bloodline of the virus.
+
In the novel ''[[Forged in Fire]]'', it is revealed that the virus did not solely affect Klingons; the ridged [[Trill]] witnessed in {{e|The Host}} – as opposed to the spotted Trill seen in ''[[Star Trek: Deep Space Nine]]'' and later episodes – are the result of a strain of the augment virus that managed to infect a Trill colony through visiting Klingon traders. However, the Trill seemed not to be interested in working on methods of restoring the original Trill look, with this "sub-group" having recently (at the time the novel was set) being re-accepted back into Trill society. The novel also reveals that the appearance of [[The Albino]] – who is not only Klingon but is in fact Kor's cousin – was an unintended side-effect of prenatal genetic engineering intended to cure his bloodline of the virus.
   
In [[Star Trek Online]], the cure to the Klingon Augment Virus is revealed. By way of the [[Guardian of Forever]], A group of Klingons travel back to [[2270]] shortly after capturing [[Miral Paris]],daughter of [[Tom Paris]] and [[B'Elanna Torres]]. At the conclusion of the mission, Miral's unqiue [[DNA]] is used to cure the Klingons of the Augment Virus.
+
In ''[[Star Trek Online]]'', the cure to the Klingon augment virus is revealed. By way of the [[Guardian of Forever]], a group of Klingons travel back to [[2270]] shortly after capturing [[Miral Paris]], daughter of [[Tom Paris]] and [[B'Elanna Torres]]. At the conclusion of the mission, Miral's unique DNA is used to cure the Klingons of the augment virus.
  +
{{bginfo|In the ''Voyager'' episode [[Prophecy (episode)|Prophecy]], a group of Klingons was cured of a virus by Tom and B'Elanna's then-unborn child, due to its unique DNA.}}
   
 
== External links ==
 
== External links ==
* [http://www.startrek.com/startrek/view/features/documentaries/article/1614.html Klingon Discrepancy Theories: Where Did the Ridges Go?], a [[Startrek.com]] documentary from {{d|22|July|2003}}.
+
*{{brokenlink|url=http://www.startrek.com/startrek/view/features/documentaries/article/1614.html|text=Klingon Discrepancy Theories: Where Did the Ridges Go?}}, a [[StarTrek.com]] documentary from {{d|22|July|2003}}
*{{Wikipedia|Retrovirus}}
+
* {{wikipedia|Retrovirus}}
*{{Wikipedia|Horizontal gene transfer}}
+
* {{wikipedia|Horizontal gene transfer}}
   
   
 
[[de:Klingonischer Augment-Virus]]
 
[[de:Klingonischer Augment-Virus]]
[[nl:Klingon augment virus]]
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[[fr:Virus amélioré klingon]]
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[[nl:Klingon augmentvirus]]
 
[[Category:Medical conditions]]
 
[[Category:Medical conditions]]

Revision as of 07:07, 4 April 2016

Klingon augment virus Levodian flu

The Klingon augment virus

"We do not discuss it with outsiders."

The Klingon augment virus was a modified form of Levodian flu that threatened to wipe out the Klingon race in the mid-22nd century. It was inadvertently created by Klingon researchers who were attempting to bio-engineer enhanced warriors using DNA from genetically-modified Human embryos left over from Earth's Eugenics Wars.

History

Klingon cranial ridges dissolve

Dissolving cranial ridges

In 2154, Klingon scientists working under the direction of Antaak recovered several Augment embryos from a hijacked Bird-of-Prey destroyed in orbit of Qu'Vat colony during the Augment Crisis. Fearing that Humans could overwhelm the Empire if they moved to staffing their ships with Augments, a project was set up to attempt to create their own Augments to counter the perceived threat.

The effort failed dreadfully. The Augments created did develop enhanced strength and intelligence, but they also began to show Human characteristics, from the blatant to the subtle. These included personality alterations and loss of the characteristic forehead ridges. Ultimately, the subjects died agonizing deaths when the incompatible DNA resulted in neural system breakdown. While working to stabilize the augmentation process, the scientific team enhanced a test subject who was, unknown to the team, also infected with Levodian flu. The flu virus in some manner incorporated the Augment DNA, turning a carefully controlled experiment into an epidemic. As the unstable augmentation process always killed the subject, this epidemic threatened the Klingon race with extinction.

The virus spread very rapidly, infecting millions within months. One of the first planets struck was N'Vak colony. The Klingons attempted to capture Doctor Arik Soong to help develop a cure, but their efforts were thwarted by Soong's incarceration in a high security detention facility. They then moved to kidnap Doctor Phlox, using Rigelian intermediaries and the assistance of Section 31 agents. Phlox was forced to work with Antaak to find a cure. However, the Klingon High Council was unwilling to wait, and dispatched a fleet to annihilate the infected planets, starting with N'Vak colony. Afterward it set course to Qu'Vat colony. (ENT: "Affliction")

Antaak

Antaak before

Antaak-ridgeless

Antaak after

When it became clear to the research team on Qu'vat colony that it would take weeks to develop a cure, stabilizing the augment virus became the researcher's primary concern. Their efforts proved successful; a method was developed that stabilized the process in the early stages, after the cranial ridges had dissolved and some minor neural re-ordering had occurred (which caused personality changes), but before the virus became contagious, enhanced augment abilities, and caused a fatal neural pathway breakdown.

Eventually, Admiral Krell convinced the High Council to call off their sterilization program of destroying the infected planets. The cure was distributed throughout the Empire. Because afflicted Klingons' DNA had been altered by the virus, the Human appearance and genetic traits were passed onto the descendants of the infected. Initially there were millions of Klingons who had to live with the disfigurement. (ENT: "Divergence")

Klingon invasion of Organia

A group of Klingons descended from those infected with the augment virus in 2267

In 2154, Antaak and Laneth feared that Klingons without ridges would become outcasts, lose their positions and would no longer have a place in the Empire. (ENT: "Divergence") However, by the 2260s, affected Klingons served as the captains and crews of front-line starships in the Klingon Defense Force. (Star Trek: The Original Series) Later on, some of them even achieved the title of Dahar Master and earned their statues among the Hall of Heroes on Qo'noS. (DS9: "Blood Oath", "The Sword of Kahless")

By the 24th century, knowledge of Klingons with smooth foreheads had been generally forgotten by the population of the United Federation of Planets. Also, the reason for Klingons having smooth foreheads a hundred years earlier was not widely known outside the Empire. When asked by Miles O'Brien and Julian Bashir, Worf simply gave the answer "we don't discuss it with outsiders", though Bashir immediately suspected viral mutation to be the cause. (DS9: "Trials and Tribble-ations")

In TOS: "The Savage Curtain", the Excalbians created Kahless as an affected Klingon. This may however be attributed to the fact that they created him from the minds of Humans who at that point mostly had dealings with smooth forehead Klingons.
A cure to restore Klingons to their proper appearance was presumably found some time between 2268 and 2293, as Kang was seen as a smooth-headed Klingon in TOS: "Day of the Dove" and as a normal Klingon in VOY: "Flashback". The first time normal Klingons were seen on screen was in Star Trek: The Motion Picture set in the early-2270s.

Background information

This storyline, seen in the Enterprise episodes "Affliction" and "Divergence", was intended in part to explain why Original Series Klingons had a more Human appearance, and later Klingons did not. The writers also hoped the two-parter would shed some light on the change in the Klingons' temperament and disposition between the TOS and TNG eras.

For years, unofficial explanations had appeared in Star Trek literature to explain the difference between the Klingons from the 1960s series and those in later productions. The idea of genetic engineering was explored heavily in several publications licensed by Paramount Pictures, among them FASA's Star Trek: The Roleplaying Game, several Pocket Books novels (such as Rules of Engagement), and the reference book The Worlds of the Federation by author Shane Johnson. These works indicated that the "Klingons" encountered during the Original Series era were "Klingon-Human fusions" intentionally created to make infiltration into Federation space easier. The true nature of Klingons was revealed during the emergency transmission of the IKS Amar as depicted in Star Trek: The Motion Picture.

In addition to the "fusion" explanation, the producers of Star Trek: Enterprise had heard many pitches over the years for stories aimed at explaining the change in the Klingon appearance. Additional theories included that the Klingons seen during the Original Series were from another race than those of later eras and also that some sort of disease was to blame. The Enterprise writing staff mirrored the fan community in that some felt it might be fun to account for the change, while others thought it was probably best to ignore the entire situation. The topic came up again when writer Manny Coto was named showrunner during the series' fourth season.

At first, the producers were only interested in a story with one or more "ridgeless" Klingons who had infiltrated Starfleet for the purpose of intelligence gathering. It was thought the story might involve a surgically-altered Klingon operative aboard the Enterprise, someone like Arne Darvin a century later. Around this time, the writing staff had recently concluded the three-part Augment Crisis arc, and it occurred to them that some of the genetically-engineered embryos might have survived the destruction of the Bird-of-Prey, and that the Klingons might use these embryos to bio-engineer their own version of "Klingon supermen". This seemed to be a way into a story dealing with the origin of Human-like Klingons. More to the point, the Enterprise producers thought it was simply "too cool" an idea to reveal that Kor, Kang, and other Original Series Klingons may have had the DNA of Khan inside them.

While some fans clamored for an explanation for the "smooth forehead" Klingons, "Affliction" teleplay writer Mike Sussman hoped to do more – account for the apparent change in Klingon culture between the 23rd and 24th centuries. As depicted in the first Star Trek series, Klingons were notoriously savage, crafty, and at times even cowardly, putting this characterization at odds with the more noble and honorable Klingons seen in The Next Generation. The "minor neural re-ordering" noted by Phlox was intended to suggest that the augmented Klingons were not only "cursed" with a more Human appearance, but that they also inherited many Human weaknesses as well. In "Divergence", the female Laneth, newly infected by the augment virus, claimed that she "felt fear for the first time since I was a child." Her fellow augmented warriors had "become like (Humans)... weak, cowardly."

The Enterprise writers' explanation for the change in the Klingons did not – and could not – satisfy every Star Trek fan. Gene Roddenberry himself reportedly believed any "explanation" was unnecessary; the makeup seen in the films and the later series would have been too expensive during the 1960s. Roddenberry felt it was best to simply imagine that Klingons always had ridges (although this preference had already been contradicted by canon when the change was noted by the DS9 crew in "Trials and Tribble-ations").

This background information was contributed in part by former Enterprise writer/producer Mike Sussman, a member of the Memory Alpha community.

Apocrypha

In The IDIC Epidemic, an original series novel written during the airing of The Next Generation, a Klingon scientist, Korsal, refers to the "Imperial Race," a reclusive breed of Klingons who retained their brow ridges. The Imperial Race has never been seen outside the Empire and its existence is largely a mystery to the Federation. The plague that is the subject of the novel is proven to be a mutated form of "The Imperial Plague," to which the Imperial race is very vulnerable, while smooth-headed Klingons are immune. This novel contained its own partial explanation for the difference in appearance between Klingons from the Original Series and The Next Generation - to wit, that "smooth-headed" Klingons were an offshoot of the Klingon race specifically engineered or selected for contact with outside forces.

In "Against Their Nature", the first installment of the IDW Comics Star Trek: Klingons - Blood Will Tell series, it is suggested that, while Phlox's cure removed Augment strength and Augment intelligence, those affected did retain Augment ambition, and as such they were largely responsible for the expansion of the Klingon Empire between Enterprise and TOS, and were able to take control of the High Council.

In the novel Forged in Fire, it is revealed that the virus did not solely affect Klingons; the ridged Trill witnessed in "The Host" – as opposed to the spotted Trill seen in Star Trek: Deep Space Nine and later episodes – are the result of a strain of the augment virus that managed to infect a Trill colony through visiting Klingon traders. However, the Trill seemed not to be interested in working on methods of restoring the original Trill look, with this "sub-group" having recently (at the time the novel was set) being re-accepted back into Trill society. The novel also reveals that the appearance of The Albino – who is not only Klingon but is in fact Kor's cousin – was an unintended side-effect of prenatal genetic engineering intended to cure his bloodline of the virus.

In Star Trek Online, the cure to the Klingon augment virus is revealed. By way of the Guardian of Forever, a group of Klingons travel back to 2270 shortly after capturing Miral Paris, daughter of Tom Paris and B'Elanna Torres. At the conclusion of the mission, Miral's unique DNA is used to cure the Klingons of the augment virus.

In the Voyager episode Prophecy, a group of Klingons was cured of a virus by Tom and B'Elanna's then-unborn child, due to its unique DNA.

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