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The Caves of Mak'ala were a series of interconnecting underground caves on Trill. Deep within these caves were kilometers of interconnecting pools of a milk-like substance which was the natural environment of the Trill symbionts.
The symbionts would die unless they were in the milk pools or were joined after a certain amount of time. When living in the pools, they could communicate with one another via electrical pulses sent through the liquid. They bred and were born in the pools.