The Caves of Mak'ala
The Caves of Mak'ala were a series of interconnecting underground caves on the Trill homeworld. Deep within these caves were kilometers of interconnecting pools of a milk-like substance which was the natural environment of the Trill symbionts.
The symbionts will die unless they are in the milk pools or are joined after a certain amount of time. When living in the pools, they can communicate with one another via electrical pulses sent through the liquid. They breed and are born in the pools.
The Guardians are a group of unjoined Trill who have devoted their lives to making sure the pools are perfect for the symbionts, having the perfect temperature and protein content.
In 2371, Commander Sisko, Doctor Bashir, and Jadzia Dax went to Trill and visited the pools looking for answers to a problem Jadzia was experiencing with the Dax symbiont. (DS9: "Equilibrium")
In 2375, Sisko suggested to Ezri Dax that she could live in the Caves taking care of the symbionts when she considered leaving Starfleet. (DS9: "Afterimage")
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